This is a very nice hold. Good puzzles, varying degrees of difficulty. The level four rooms were cute, although not particularly challenging and didn't seem much in keeping with the spirit of the rest of the hold.
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It may be too easy this way, but your 'queens' don't behave like roach queens, where they only spawn on empty squares.
They also don't flee, but that seems less confusing -- the spawning underneath existing creatures gets kind of weird.
Several tough puzzles here.
Overall, I give it an 8 with 6.5 brains of difficulty.
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"
Rings and knots of joy and grief, all interlaced and locking."
--William Buck